The life cycle of freshwater mussels includes a larval stage (the glochidium) which is an obligate parasite on fish gills or fins. Glochidia are retained in the gills of the female prior to release of the parasitic stage. Species vary in the length of the breeding season as well as in growth rates and shell dimorphism. Students have examined life history from an evolutionary perspective to determine the adaptive significance of alternative breeding strategies.

A second area of research deals with systematics and ecology of non-insectan terrestrial arthropods. This work focuses on systematics and biogeography of woodland centipedes, especially the Lithobiomorpha.